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OF FALCONS
A falcon is any species of raptor
in the genus Falco.
The word comes from their Latin name
falco,
related to Latin falx
("sickle") because of the shape of these birds' wings.
Overview
Adult falcons have thin tapered
wings, which enable them to fly at high speed and to change
direction rapidly. Fledgling falcons, in their first year of
flying, have longer flight feathers which makes their
configuration more like that of a general-purpose bird such
as a broadwing.
This is to make it easier for them to fly while learning the
exceptional skills required to be effective hunters in their
adult configuration. A falcon's wings are shaped like a
scythe. Common misconceptions of the difference of a scythe
and sickle are the cause of the misconception of the shape
of the falcons wings.
Peregrine Falcons have been
recorded diving at speeds of 124 miles per hour (200 km/hr),
making them the fastest-moving creatures on Earth.
Other falcons include the Gyrfalcon, Lanner Falcon, and the
Merlin. Some small falcons with long narrow wings are called
hobbies, and some which hover while hunting are called
kestrels. The falcons are part of the family Falconidae,
which also includes the caracaras, Laughing Falcon, forest
falcons, and falconets.
The traditional term for a male
falcon is tercel
(British spelling) or tiercel
(American spelling), from Latin
tertius =
third because of the belief that only one in three eggs
hatched a male bird. Some
sources give the etymology as deriving from the fact that a
male falcon is approximately one third smaller than the
female (Old French tiercelet).
A falcon chick, especially one
reared for falconry, that is still in its downy stage is
known as an eyas
(sometimes spelt eyass).
The word arose by mistaken division of Old French
un niais,
from Latin presumed *nidiscus
("nestling", from nidus
= nest). The technique of hunting with trained captive birds
of prey is known as falconry.
As is the case with many birds of
prey, falcons are renowned for their exceptional powers of
vision; one species has been found to have a visual acuity
of 2.6 times that of a normal human.
In February 2005, the Canadian
ornithologist
Louis Lefebvre announced a method of
measuring avian intelligence in terms of their innovation in
feeding habits. The falcon and
corvids scored highest on this
scale.
Systematics and evolution
Compared to other birds of prey,
the fossil record of the falcons is not well distributed in
time. The oldest fossils tentatively assigned to this genus
are from the Late Miocene, less than 10 million years ago.
This coincides with a period in which many modern genera of
birds became recognizable in the fossil record. The falcon
lineage may however be somewhat older than this
and given the distribution of fossil and living
Falco
taxa is probably of North
American, African or possibly Middle Eastern or European in
origin.
Falcons are roughly divisible into
three or four groups. The first contains the kestrels
(probably excepting the American Kestrel); usually small and
stocky falcons of mainly brown upperside color and sometimes
sexually dimorphic; three African species that are generally
grey in color stand apart from the typical members of this
group. Kestrels feed chiefly on terrestrial
vertebrates and invertebrates of
appropriate size, such as rodents, reptiles, or insects.
The second group contains slightly
larger (on average) and more elegant species, the hobbies
and relatives. These birds are characterized by considerable
amounts of dark slaty grey in their plumage; the malar area
is nearly always black. They feed mainly on smaller birds.
Third are the Peregrine Falcon and
its relatives: large powerful birds which also have a black
malar area (except some very light color morphs), and often
a black cap also. Otherwise, they are somewhat intermediate
between the other groups, being chiefly medium grey with
some lighter or brownish colours on the upper side. They are
on average more delicately patterned than the hobbies and if
the hierofalcons are excluded (see below), this group
contains typically species with horizontal barring on the
underside. As opposed to the other groups, where tail colour
varies much in general but little according to evolutionary
relatedness, the tails of the
large falcons are quite uniformly dark grey with rather
inconspicuous black banding and small white tips, though
this is probably plesiomorphic.
These large Falco
feed on mid-sized birds and terrestrial vertebrates.
Very similar to these and sometimes
included therein are the 4 or so species of hierofalcons
(literally, "hawk-falcons"). They represent taxa with
usually more phaeomelanins
which impart reddish or brown colors, and generally more
strongly patterned plumage reminiscent of hawks. Notably,
their undersides have a lengthwise pattern of blotches,
lines or arrowhead marks.
While these three or four groups,
loosely circumscribed, are an informal arrangement, they
probably contain several distinct clades in
their entirety. A study of
mtDNA
cytochrome
b
sequence data of some kestrels (Groombridge
et al.
2002) identified a clade containing the Common Kestrel and
related "malar-striped"
species, to the exclusion of such taxa as the Greater
Kestrel (which lacks a malar stripe), the Lesser Kestrel
(which is very similar to the Common but also has no malar
stripe), and the American Kestrel. The latter species has a
malar stripe, but its color pattern - apart from the
brownish back - and notably also the black feathers behind
the ear, which never occur in the true kestrels, are more
reminiscent of some hobbies. The malar-striped kestrels
apparently split from their relatives in the Gelasian,
roughly 2.5-2 mya,
and are apparently of tropical East African origin. The
entire "true kestrel" group - excluding the American species
- is probably a distinct and quite young clade, as
also suggested by their numerous apomorphies.
Other studies
have confirmed that the hierofalcons are a monophyletic
group - and, incidentally, that hybridization is quite
frequent at least in the larger species falcon species.
Initial studies of mtDNA
cytochrome
b
sequence data suggested that the hierofalcons are basal
among living falcons. This is
now known to be an erroneous result due to the presence of a
numt
(Wink & Sauer-Gürth 2000); in reality the hierofalcons are a
rather young group, originating maybe at the same time as
the start of the main kestrel radiaton, about 2 million
years ago. This lineage seems to have gone nearly extinct at
some point in the past; the present diversity is of very
recent origin, though little is known about their fossil
history (Nittinger et al.
2005, Johnson et al.
2007).
The phylogeny and delimitations of
the Peregrine and hobbies groups is more problematic.
Molecular studies have only been conducted on a few species,
and namely the morphologically ambiguous taxa have often
been little researched. The morphology of the syrinx,
which contributes well to resolving the overall phylogeny of
the Falconidae, is not very informative in the present
genus. Nonetheless, a core group containing the Peregrine
and Barbary falcons which in turn group with the
hierofalcons and the more distant Prairie Falcon (which was
sometimes placed with the hierofalcons, even though it is
entirely distinct biogeographically), as well as at least
most of the "typical" hobbies, are confirmed to be
monophyletic as suspected.
Given that the American
Falcos of
today belong to the Peregrine group or are apparently more
basal species, it seems that the initially most successful
evolutionary radiation was an
Holarctic one that originated
possibly around central Eurasia or in (northern) Africa. One
or several lineages were present in North America by the
Early Pliocene at latest.
In conclusion, the origin of
today's major Falco
groups - the "typical" hobbies and kestrels for example, or
the Peregine-hierofalcon complex, or the Aplomado Falcon
lineage - can be quite confidently placed from the Miocene-Pliocene
boundary through the Zanclean
and Piacenzian
and just into the Gelasian, that is from about 8 to 2.4
million years ago, when the malar-striped kestrels
diversified. Some groups of falcons, such as the hierofalcon
complex or the Peregrine-Barbary superspecies have only
evolved in more recent times; the species of the former seem
to be a mere 120.000 years old or so (Nittinger
et al.
2005).
At Falconry-UK we have
three Falcons.
All are Parent Reared and
we have had since birth.
Please click on the
Species Name below to view photographs or the Name of Falcon
to view them individually.
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